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Preliminary experiments demonstrated (a) that the light doses used were saturating for phytochrome photoconversions and (b) that the centrifugation procedure adopted pelleted the same amount of phytochrome as the 20,000 x g for 30 min commonly used in the literature. Phytochrome measurements. Phytochrome was assayed as previously described with a modified (Marme, 1969) Ratiospect using C a C 0 as a scattering agent (Butler and 3 Norris, 1960). "Percent pelletable phytochrome" represents the amount in the 50,000 x g pellet as a percentage of the combined total of supernatant and pellet.

Tissue irradiated and incubated at 0°C was held bathed in extraction medium in tubes on ice during both operations. Tissue irradiated and incubated at 25°C was held in tubes in a water bath at that temperature and rapidly cooled at the times indicated by addition of ice-cold extraction medium and transfer of the tubes to ice. Pre-chilled tissue (2 g) was homogenized for 5 s in 6 mi of ice-cold extraction medium using an Ultra Turrax blender at 6700 rpm and the homogenate filtered through nylon cloth.

Shoot tissue was irradiated at 25°C as in+ addition of ice-cold extraction dicated and chilled 2 by medium (minus M g ) either immediately (where the terminal irradiation was far red) or after 3 min dark incubation (where the terminal irradiation was red). After a further 3 min on ice the tissue was homogenized for 5 s. 2+ given immediThe 5 s in vitro red irradiation indicated was ately following the homogenization. M g was added to the crude homogenates at the times indicated to a final concentration of 10 mM.

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